Ophioglossaceae: Botrychium ssp. (grape ferns & moonworts )
20170923 Decided they are mostly bipinnate, so that would lead to Botrychium biternatum. One is much more dissected and could be Botrychium dissectm. Transplanted the B dissect from the edge of the driveway to copse 1 near ebony spleenwort and other Botrychium biternatum. Transplanted Botrychium biternatum to courtyard.
20161123 currently think the many moonworts observed are B dissectum
GENUS: Botrychium
NTP Species search
http://www.discoverlife.org/mp/20q?guide=Ferns : lists 16 Ophioglossaceae in North Carolina, including 6 Ophioglossum spp. Has own collection of images.
Fryer, Janet L. 2014. Botrychium spp. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/plants/fern/botspp/all.html [2017, September 23].
Radford, A. E., H. E. Ahles & C. R. Bell. 1968. Man. Vasc. Fl. Carolinas i–lxi, 1–1183. University of North Carolina Press, Chapel Hill - distinguishes between B biternatum & B dissectum as bipinnate & tripinnate, respectively.
Species in North Carolina per PLANTS
** Botrychium biternatum (Sav.) Underw. (sparselobe grapefern) published Botanical Gazette 22(5): 407. 1896
Plants: BOBI in Chatham (no images)Go Botany key X; Discover life (images)eFlora; Botrychium biternatum often grows with B . dissectum and B . jenmanii . The name B . biternatum was misapplied by L.Underwood to B . lunarioides (W.H. Wagner Jr. 1961).Radford, A. E., H. E. Ahles & C. R. Bell. 1968. Man. Vasc. Fl. Carolinas. University of North Carolina Press, Chapel Hill [kindle edition] (Shows in chatham)
** Botrychium dissectum Spreng. (cutleaf grapefern) published Anleitung zur Kenntniss der Gewächse 3: 172. 1804. (Anleit. Kenntn. Gew.)
Plants: BODI2 in Chatham (images)
Go Botany key; Discover life (images);eFlora: highly variable, even within the same population.Tropicos lists many subs & variants, herbarium specimenRadford, A. E., H. E. Ahles & C. R. Bell. 1968. Man. Vasc. Fl. Carolinas. University of North Carolina Press, Chapel Hill [kindle edition] (Shows in chatham)
** Botrychium jenmanii Underw. (Dixie grapefern)
Go Botany key X; NTP XeFloras.org has description: ... occurs in a variety of habitats. In hardwoods and especially pine woods, it is associated with B . biternatum ; in open grassy places and lawns it is found with B . lunarioides.
Radford, A. E., H. E. Ahles & C. R. Bell. 1968. Man. Vasc. Fl. Carolinas. University of North Carolina Press, Chapel Hill [kindle edition] ( as B alabamense )
Botrychium lanceolatum (S.G. Gmel.) Angstr. (lanceleaf grapefern)
Plants: BOLA NC native, no county data (images)Go Botany key X; Discover life (images); NTP XeFloras.org lists two subspecies, neither in NC "Leaves usually drying up in midsummer together with other associated species. Mainly open fields" and "Leaves appearing in late spring or early summer, releasing spores later than most associated species, and dying as late as October. Mainly in shaded woods;"Clowderwood moonworts appear in late summer, thus unlikely to be B lanceolatum
Botrychium lunarioides (Michx.) Sw. (winter grapefern)
Plants: BOLU2 NC native, no county data (x images)
[Plants maps if not in chatham or broadly NC]Go Botany key X; Discover life (images);NTP: Rare Look for it in old fields, pastures, young forests, granitic flatrocks, juniper-oak-blue ash woodlands over limestone, per Weakley's FloraeFlora:Another peculiarity of this species is the tendency for the sporophores to remain curled in late fall and early winter and to become erect in February. Botrychium lunarioides is often associated with Schizachyrium scoparius Michaux and Ophioglossum crotalophoroides WalterClowderwood habitat is acidic mafic and moist; fronds appear in late summer, thus unlikely to be B lundariodes.Radford, A. E., H. E. Ahles & C. R. Bell. 1968. Man. Vasc. Fl. Carolinas. University of North Carolina Press, Chapel Hill [kindle edition]
Botrychium matricariifolium (A. Braun ex Dowell) A. Braun ex W.D.J. Koch (matricary grapefern)
Plants: BOMA2 NC native, no county data (images)
eFlora: Leaves appearing in spring, dying in late summer.Clowderwood moonworts appear in late summer, thus unlikely to be B matricariifolium
Moonworts in the news Dr. Evelyn Williams August 18, 2016
Botrychium matricariifolium Daisy-leaved moonwort
Botrychium multifidum (S.G. Gmel.) Trevis. (Leathery grapefern)
Plants: BOMU NC native, no county data (images)
Go Botany key; Discover life (images); NTP XeFlora: Leaves green over winter, appearing in spring. Widespread mainly in fieldsClowderwood moonworts appear in late summer, thus unlikely to be B multifidum
Botrychium oneidense (Gilbert) House (bluntlobe grapefern, Oneida grape fern)
Plants: BOON NC native, in three counties (X images)
Go Botany key; Discover life (images); NTP XeFlora: Leaves green over winter, sporophores seasonal, new leaves appearing in spring. In moist, shady, acidic woods and swamps; ..Botrychium oneidense commonly occurs with B . dissectum and B . multifidum . Young individuals of both may resemble B . oneidense (W.H. Wagner Jr. 1961b).Clowderwood moonworts appear in late summer, thus unlikely to be B oneidenseRadford, A. E., H. E. Ahles & C. R. Bell. 1968. Man. Vasc. Fl. Carolinas. University of North Carolina Press, Chapel Hill [kindle edition]
Botrychium simplex E. Hitchc. (little grapefern)
Plants: BOSI NC native, no county data (images)
Go Botany key; Discover life (images);NTP: NC mountains, Look for it in disturbed gravelly areas in spruce-fir forests, northern hardwoods forests, grassy balds, per Weakley's FloraeFlora: habitat/coocurranceClowderwood moonworts seem different from diagram, habitat incorrectr, thus unlikely to be B simplex.
Botrychium virginianum (L.) Sw. {rattlesnake fern)
Plants: BOVI in Chatham (images)
Go Botany key; Discover life (images);NTP: Common, Look for it in a wide range of fairly dry, mesic, and wet forests, cove forests, especially in nutrient-rich, moist bottomlands and slopes, per Weakley's FloraeFlora: Leaves seasonal, appearing in early spring and dying in late summer. Common to abundant, especially in shaded forests and shrubby second growth…Clowderwood moonworts appear in late summer, thus unlikely to be B virginianum.Radford, A. E., H. E. Ahles & C. R. Bell. 1968. Man. Vasc. Fl. Carolinas. University of North Carolina Press, Chapel Hill [kindle edition] (Shows in chatham)
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